Stable Isotope Formulas
delta = ((R_sample / R_standard) - 1) * 1000
R_sample = R_standard * (1 + delta/1000)
R_VPDB (13C/12C) = 0.011237
R_AIR (15N/14N) = 0.003676
Trophic Level ~ (d15N - baseline) / 3.4 + 1
Stable isotopes of carbon distinguish C3 plants (d13C ~ -26 per mil) from C4 plants like maize (d13C ~ -12 per mil). Nitrogen-15 increases ~3.4 per mil per trophic level. Marine protein consumers show enriched d13C and d15N values compared to terrestrial diets. Acceptable C:N ratios for collagen are 2.9-3.6.